TRANSPORTATION OF MINERALS OF LIVING THINGS.
The Concept of Transportation of Materials in Living Things
The Functions of Root Hairs in Absorption and Movement of Water and Mineral Salts in Plants
Explain the functions of root hairs in absorption and movement of water and mineral salts in plants
Water and mineral uptake by roots
Plants absorb water from the soil through the root and transport it to the stem, leaves andflowers. Roots have root hairs that are unicellular, thin-walled outgrowths of the epiblema (skinof the root).
The root hairs are in close contact with the thin film of water surrounding the soil particles.There are mineral salts such as nitrates, chlorides, sulphates, phosphates, etc., dissolved in thiswater.
Water is absorbed by osmosis, while the minerals are absorbed as ions by active transport(transport against the law of diffusion, by spending cellular energy). The cell membrane hastransport proteins that allow the ions to cross the membrane. The ions then move upward throughthe xylem, to the leaves and other aerial parts of the plant.
The cell wall of each root hair is permeable to water and minerals, but its cell membrane and themembrane around the vacuole are semi permeable membranes. The root hair cells take upmineral ions by active transport.
This creates a concentration difference of these ions between the root and the soil. Now, the soilsolution has higher water content than the cell sap of the root hair. Hence, water from the soildiffuses into the root hair. The root hair cells now become turgid, while the adjacent cells of thecortex have lower water content.
This results in the diffusion of water from the root hairs into the cortical cells (see figure below).After passing through the cortical cells by osmosis, the water reaches the endodermis (tissueseparating the cortex from the vascular tissues). The endodermis forces water into the xylemtubes through passage cells.
The pressure with which water is pushed into the xylem tubes of the root is called root pressure.The water moving upwards forms a column, which is maintained up to a certain height due toroot pressure.
The Movement of Water and Dissolved Mineral Salts in Plants
Outline the movement of water and dissolved mineral salts in plants
Upward movement of water within the plant
There are several processes that enable the water to move up a plant. These processes includeroot pressure, transpiration pull, cohesion, adhesion and capillarity.
Root pressure
As long as the soil is damp, there will be water taken in by the root hairs. As more water is takenin, the water that is already in the xylem vessel will be pushed up the plant. This is called rootpush or root pressure and it helps to push water up to the leaves.
Root pressure is capable, under ideal atmospheric conditions, of pushing water one or two feetabove the ground. Since root pressure is not strong enough to move water up very high, anotherprocess called transpiration pull is needed to enable the water to continue moving up the plant.
Experiments to Demonstrate Transpiration pull, Root Pressure and Capillarity
Conduct experiments to demonstrate transpiration pull, root pressure and capillarity
Transpiration pull
Transpiration is the loss of water through the leaves and other parts of the plant. Mosttranspiration occurs through openings, called stomata, on the underside of the leaves. Astranspiration occurs, water is lost. This water is replaced by water in the xylem vessels. Thiscauses an upward pull (transpiration pull or transpiration stream) on the water in the vessels.Thus, water is pulled up through the plant, and more enters by the roots to replace it.
Cohesion
Cohesion is the force of attraction between similar molecules. Transpiration pull is possiblebecause water molecules cling to each other by cohesion. When water molecules cling to eachother as they move up the stem and into the leaves, they pull up more water molecules up theplant. This process, however, is facilitated by transpiration pull since the water molecules lostthrough transpiration is being replaced by more water molecules absorbed by the roots.
Adhesion
Adhesion
Adhesion is the force of attraction between different molecules. As water molecules are stucktogether by cohesion, the entire column of water in the xylem adheres to the sides of the xylem.It is said that the water in under tension as the column moves up the xylem. At the same time, thexylem tube narrows because of the tension.
Cohesion and adhesion forces maintain a continuous column of water in the xylem vessels fromthe roots to the leaves of plants.
Capillarity
Capillarity is the tendency of water to rise through narrow tubes. The lumen of xylem tracheidsand vessels is very narrow and this enables water to rise through it by capillarity. Capillarity isassisted by adhesion and cohesion forces.
The Concept of Transpiration
Explain the concept of transpiration
Transpiration is the evaporation of water from plants. It occurs chiefly through the leaves whiletheir stomata are open for the passage of carbon dioxide and oxygen during photosynthesis.
Transpiration also occurs through the cuticle and lenticels. Lenticels are pores in the stems ofwoody plants that allow gaseous exchange between the atmosphere and the internal tissues.
The Significance of Transpiration in Plants
Outline the significance of transpiration in plants
Transpiration is of immense importance in plant life as it is of great benefit to the plant. Thefollowing are the reasons why transpiration is important in plants.
Cooling of the plant
The leaves absorb the radiant energy. Some of the light energy is utilized in photosynthesis. Therest is converted into heat energy resulting in an increase in leaf temperature. However, rapidloss of water in the form of water vapour from the aerial parts of the plant through transpirationbrings down their temperature. Transpiration thus provides a significant cooling effect whichkeeps the plant from being overheated.
Mineral transport
Mineral salts remain dissolved in the soil water and are absorbed by the roots. Minerals that arcabsorbed and accumulated in the xylem duct of the root move up and are distributed in the plantby the transpiration stream.
Water movement
The absorbed water is transported from roots to leaves through the xylem vessels. This is greatlyinfluenced by transpiration pull. Water loss due to transpiration results in the development of lowwater potential in the leaf tissues. Thus, water moves from the xylem vessels to the leaf cells.
Development of mechanical tissues
Greater amount of transpiration helps in the development of mechanical tissues in plants. Theplants become healthier and more compact, the cell walls become thick and cutinized and theplants are able to resist the attack of fungi and bacteria.
Maintenance of turgidity
Transpiration maintains an optimum degree of turgidity in cells. Under favourable conditions,plants absorb excess amount of water, which is given off by transpiration to maintain theoptimum turgidity for better growth.
Increase of taste of fruits
The solutes inside the cell become more concentrated when transpiration is rapid. Consequently,the concentration of sugar solution in the cells of fruits increases and fruits taste sweeter.
Wilting
When the rate of evaporation is higher than that of absorption of water from the soil, as it occursduring drought conditions, the plant wilts. Wilting is beneficial when a plant cannot obtainenough water to replace that lost by the plant through transpiration because it causes the closureof the stomata (singular: stoma). Thus, the rate of evaporation is greatly reduced.
Transpiration as a necessary evil
Transpiration is a necessary evil because of the following facts:-
- A large amount of absorbed water is lost during transpiration which is harmful to plants.
- Unnecessary wastage of energy takes place during the process of water absorption which islost due to transpiration.
- When the rate of transpiration is high in plants growing in soil deficient in water, an internalwater deficit develops in plants which may affect metabolic process.
- Many xerophytes undergo structural modifications and adaptations to check transpiration.
Considering both the beneficial and harmful effects of transpiration, it may be concluded that itis definitely advantageous in spite of its harmful consequences.
Factors Affecting the Rate of Transpiration in Plants
Outline factors affecting the rate of transpiration in plants
The rate of transpiration can be affected by both plant features and environmental factors.
Plant factors
These plant parameters help plants control rates of transpiration by serving as forms of resistanceto water movement out of the plant. They include the following:-
Root system
Plants with extensive root systems absorb a great amount of water and therefore much water ismoved up the plant. Thus, plants with extensive root systems have higher rates of transpirationthan those with few roots.
Size of leaves
A plant with broad leaves tend to lose more water than that with small leaves. This is because thebroad leaves have large surface areas over which transpiration takes place.
Leaf structure
The structure of a leaf has a great influence on the rate of transpiration. The following areanatomical structures of a leaf that affect the rate of transpiration:-
Number of stomata
Stomata are pores in the leaf that allow gaseous exchange to take place, and water vapour toleave the plant. Special cells called guard cells control each pore’s opening or closing. Someplants have many stomata while others have a few stomata. The more the stomata, the higher therate of transpiration and vice versa.
Position of stomata
Plants with few stomata on the upper surface of the leaf experiences a little transpirationcompared to those with many stomata on the lower leaf surface. This is because the uppersurface is highly stricken by direct sunlight hence increasing the rate of transpiration.
Epidermal hairs
Epidermal hair on the leaf traps a thin layer of still air close to the leaf surface. For the water lostfrom the leaf to get into the atmosphere, it has to cross this resistant layer of air. The layer thuschecks excessive loss of water from the leaf. Likewise, the water vapour from the leaf is alsotrapped by the epidermal hairs. This prevents further loss of water vapour from the leaves andhence slows down the rate of transpiration.
Size of stomatal air spaces
Large air spaces between the cells of the spongy mesophyll and stomata, called substomatal airspaces, increase the rate of transpiration. Small substomatal air spaces reduce the rate oftranspiration.
Cuticle
The cuticle is the waxy layer present on all above-ground tissue of a plant and serves as a barrierto water movement out of a leaf. Because the cuticle is made of wax, it is very hydrophobic or‘water-repelling’. Therefore, water does not move through it very easily. The thicker the cuticlelayer on a leaf surface, the slower the transpiration rate. Cuticle thickness varies widely amongplant species. In general, plants from hot, dry climates have thicker cuticles than plants fromcool, moist climates. In addition, leaves that develop under direct sunlight will have muchthicker cuticles than leaves that develop under shade conditions.
Environmental factors
Some environmental conditions create the driving force for movement of water out of the plant.Others alter the plant’s ability to control water loss.
Light
Plants transpire more rapidly in the light than in the dark. This is largely because light stimulatesthe opening of the stomata (mechanism). Light also speeds up transpiration by warming the leaf.
Photosynthesis occurs in the presence of light. A higher light intensity increases the rate ofphotosynthesis in the guard cells. As the guard cells absorb water from the soil forphotosynthesis, they become turgid and hence the stomata are opened, and hence a higher rate oftranspiration.
Temperature
Plants transpire more rapidly at higher temperatures because water evaporates more rapidly asthe temperature rises. At 30°C, a leaf may transpire three times as fast as it does at 20°C.
Relative humidity
Relative humidity is the amount of water vapour in the air compared to the amount of watervapour that air could hold at a given temperature. When the air is less moist, the relativehumidity is low, and thus the rate of transpiration is greater. When relative humidity is high, theatmosphere contains more moisture, reducing the rate of transpiration. Therefore, transpirationincreases with the decrease in relative humidity.
The rate of diffusion of any substance increases as the difference in concentration of thesubstances in the two regions increases. When the surrounding air is less humidity, diffusion ofwater out of the leaf goes on more rapidly.
Wind
The wind removes water vapour and thus increases the rate of transpiration. High winds lead tostomatal closure to stop the rapid water loss and hence bring a drop in rate of transpiration.Moderate winds may reduce transpiration by lowering the temperature of the leaf.
When there is no breeze, the air surrounding a leaf becomes increasingly humid thus reducingthe rate of transpiration. When a breeze is present, the humid air is carried away and replaced bydrier air, thus increasing the rate of transpiration.
Soil water
The source of water for transpiration out of the plant comes from the soil. Plants with adequatesoil moisture will normally transpire at high rates because the soil provides the water to movethrough the plant. Plants cannot continue to transpire without wilting if the soil is very drybecause the water in the xylem that moves out through the leaves is not being replaced by the soil water. Thus, the rate of transpiration will increase when there is adequate amount of water inthe soil and will decrease when the soil contains little mositure.
Atmospheric pressure
Transpiration is high at low atmospheric pressure and it is low at high atmospheric pressure.Plants that grow naturally at higher altitudes, where the atmospheric pressure is low, havemodified leaves to reduce the rate of transpiration.